- Physical interaction of Epstein-Barr virus (EBV) nuclear antigen leader protein (EBNA-LP) with human oestrogen-related receptor 1 (hERR1): hERR1 interacts with a conserved domain of EBNA-LP that is critical for EBV-induced B-cell immortalization. Igarashi, M., Kawaguchi, Y., Hirai, K., Mizuno, F. J. Gen. Virol. (2003)