- Mutations of the X-linked genes encoding neuroligins NLGN3 and NLGN4 are associated with autism. Jamain, S., Quach, H., Betancur, C., Råstam, M., Colineaux, C., Gillberg, I.C., Soderstrom, H., Giros, B., Leboyer, M., Gillberg, C., Bourgeron, T. Nat. Genet. (2003)