- The genetic heritage of the earliest settlers persists both in Indian tribal and caste populations. Kivisild, T., Rootsi, S., Metspalu, M., Mastana, S., Kaldma, K., Parik, J., Metspalu, E., Adojaan, M., Tolk, H.V., Stepanov, V., Gölge, M., Usanga, E., Papiha, S.S., Cinnioğlu, C., King, R., Cavalli-Sforza, L., Underhill, P.A., Villems, R. Am. J. Hum. Genet. (2003)