- Lipophilic quaternary ammonium salt acts as a mucosal adjuvant when co-administered by the nasal route with vaccine antigens. Klinguer, C., Beck, A., De-Lys, P., Bussat, M.C., Blaecke, A., Derouet, F., Bonnefoy, J.Y., Nguyen, T.N., Corvaïa, N., Velin, D. Vaccine (2001)