- Fragmin60 encodes an actin-binding protein with a C2 domain and controls actin Thr-203 phosphorylation in Physarum plasmodia and sclerotia. Sklyarova, T., De Corte, V., Meerschaert, K., Devriendt, L., Vanloo, B., Bailey, J., Cook, L.J., Goethals, M., Van Damme, J., Puype, M., Vandekerckhove, J., Gettemans, J. J. Biol. Chem. (2002)