- The upstream region of the isocitrate lyase gene (UPR-ICL) of Candida tropicalis induces gene expression in both Saccharomyces cerevisiae and Escherichia coli by acetate via two distinct promoters. Atomi, H., Umemura, K., Higashijima, T., Kanai, T., Yotsumoto, Y., Teranishi, Y., Ueda, M., Tanaka, A. Arch. Microbiol. (1995)