- The Werner syndrome helicase and exonuclease cooperate to resolve telomeric D loops in a manner regulated by TRF1 and TRF2. Opresko, P.L., Otterlei, M., Graakjaer, J., Bruheim, P., Dawut, L., Kølvraa, S., May, A., Seidman, M.M., Bohr, V.A. Mol. Cell (2004)