- Arabidopsis VARIEGATED 3 encodes a chloroplast-targeted, zinc-finger protein required for chloroplast and palisade cell development. Naested, H., Holm, A., Jenkins, T., Nielsen, H.B., Harris, C.A., Beale, M.H., Andersen, M., Mant, A., Scheller, H., Camara, B., Mattsson, O., Mundy, J. J. Cell. Sci. (2004)