- Drosophila dSAP18 is a nuclear protein that associates with chromosomes and the nuclear matrix, and interacts with pinin, a protein factor involved in RNA splicing. Costa, E., Canudas, S., Garcia-Bassets, I., Pérez, S., Fernández, I., Giralt, E., Azorín, F., Espinás, M.L. Chromosome Res. (2006)