- A SPGY copy homologous to the mouse gene Dazla and the Drosophila gene boule is autosomal and expressed only in the human male gonad. Shan, Z., Hirschmann, P., Seebacher, T., Edelmann, A., Jauch, A., Morell, J., Urbitsch, P., Vogt, P.H. Hum. Mol. Genet. (1996)