- LPP, an actin cytoskeleton protein related to zyxin, harbors a nuclear export signal and transcriptional activation capacity. Petit, M.M., Fradelizi, J., Golsteyn, R.M., Ayoubi, T.A., Menichi, B., Louvard, D., Van de Ven, W.J., Friederich, E. Mol. Biol. Cell (2000)