- alphadbeta2 integrin is expressed on human eosinophils and functions as an alternative ligand for vascular cell adhesion molecule 1 (VCAM-1). Grayson, M.H., Van der Vieren, M., Sterbinsky, S.A., Michael Gallatin, W., Hoffman, P.A., Staunton, D.E., Bochner, B.S. J. Exp. Med. (1998)