- Betaglycan expression is transcriptionally up-regulated during skeletal muscle differentiation. Cloning of murine betaglycan gene promoter and its modulation by MyoD, retinoic acid, and transforming growth factor-beta. Lopez-Casillas, F., Riquelme, C., Perez-Kato, Y., Ponce-Castaneda, M.V., Osses, N., Esparza-Lopez, J., Gonzalez-Nunez, G., Cabello-Verrugio, C., Mendoza, V., Troncoso, V., Brandan, E. J. Biol. Chem. (2003)